1- Androschchuk A. F. and Kostinenko L. D., (1981) Chromosome numbers of the genus Achillea L. Certain species cultivated in botanical gardens. Ukrajinsk Bot. Zurn. (Kiev), 38: 53-57.
2- Benedek B. and Kopp B. (2007) Achillea millefolium L. s. l. revisited: recent findings confirm the traditional use. Wien Med Wochenschr. 157: 312-314
3- Bremer K. and Humphries C. J. (1993) Generic monograph of the Asteraceae-Anthemideae. Bull. Nat. Hist. Mus. London (Bot.) 23: 71-177.
4- Camacho J. P. M., Sharbel T. F., and Beukeboon L. W. (2000) B-chromosome evolution. Philos Trans R Soc Lond B Biol Sci 355:163–178
5- Carr G. D., King R. M., Powell A. M. and Robinson H. (1999) Chromosome number in Compositeae XVIII. Am. Jour. Bot. 86: 1003-1013.
6- Chehregani A. and Mehanfar N. (2008) New Chromosome Counts in the Tribe Anthemideae (Asteraceae) from Iran. Cytologia 73: 189-196.
7- Chehregani A. and Hajisadeghian S. (2009) New chromosome counts in some species of Asteraceae from Iran. Nordic Jour. Bot. 27: 247-250.
8- Chehregani A., Atri M., Yousefi S. and Jalali F. (2010) Polyploidy variation in some species of the genus Artemisia L. (Asteraceae) in Iran. Caryologia 63: 168-175.
9- Chehregani A., Mohsenzaeh F., Moazen F. (2012) Variation of chromosome numbers in Tanacetum parthenium Schultz Bip. (Asteraceae) in Iran. Chromosome Botany 7: 97-100.
10- Chehregani A., Heidarian M., Jalali F. (2013) Cytological investigations on Achillea wilhelmsii C. Koch (Asteraceae) in the western part of Iran. Chromosome Botany, 8:13-16.
11- Contandriopoulos J. and Martin D., (1967) Contribution a I’etude cytotaxonomique des Achillea de Grece. Irregularite de la meiose. Bull. Bot. Soc. France, 114: 257-275.
12- Constantinidis T. H. and Kalpotzakis E., (2005) A new species of Achillea L. (Asteraceae: Anthemideae) from South-East Peloponnisos, Greece. Bot. J.Linn. Soc. 147: 249–256.
13- D˛abrowska J., 1989. The chromosome number of several taxa of the genus Achillea L. Acta Soc. Bot. Poloniae 58: 163-177.
14- D˛abrowska J. (1992) The chromosome number and DNA content in taxa of Achillea L. In relation to the distribution of the genus. Acta Universitatis Wratislaviensis No. 1425, Prace Botaniczne 49: 1–84.
15- Danihelka J. and Rotreklov O., (2001) Chromosome numbers within the Achillea millefolium and the A. distans groups in the Czech Republic and Slovakia. Folia Geobot. 36: 163–191.
16- Danihelka J. and Rotreklov O., (2002) Achillea distans (Asteraceae) confirmed as native in the Bieszcady Mts (SE Poland). Fragmenta Floristica et Geobotanica 45 (2000): 257–263.
17- Dyer A. F., Jong K., Ratter J. A. (1970) Aneuploidy: a redefinition. Notes from the Royal Botanic Garden Edinburgh 30: 177-182.
18- Efimov K. F., (1998) Chromosome numbers in some species of the families Asteraceae, Campanulaceae, Fabaceae from central Caucasus. Bot. Žurn. (Moscow & Leningrad) 83(8): 131–132.
19- Efimov K. F., (2005) Chromosome numbers of some species of the families Amaryllidaceae, Asteraceae, Fabaceae in North Osetia. Pages 33-35 in Karyology, Karyosystematics and Molecular Phylogeny. St. Petersburg, Russia.
20- Ehrendorefer F., and Guo Y.P. (2006) Multidisciplinary studies on Achillea sensu lato (Compositae –Antemideae ): new data on systematic and phylogeography Willdenowia 36: 69-87.
21- Funk V. A., Susanna A., Steussy T. F., and Bayer R. J. (2009) Systematics, evolution and biogeography of Compositae, International Association for Plant Taxonomy.
22- Ghaffari S. M. and Kelich K. (2006) New or rare chromosome counts of some angiosperm species from Iran. Iran. Jour. Bot. 12: 81-86.
23- Goli S. A. H., Rahimmalek M., Sayed Tabatabaei B. E. (2008) Characteristics and fatty acid profile of yarrow (Achillea tenuifolia) seed oil. Int. J. Agric. Biol. 10, 355–357.
24- Guo Y. P., Ehrendorfer F., and Samuel R. (2004) Phylogeny and systematics of Achillea (Asteraceae-Antemideae ) inferred from nrITS and plastid trnL-F DNA sequences. Taxon 53: 657-672
25- Guo Y. P., Saukel J., Mittermayr R., Ehrendorfer F. (2005) AFLP analyses demonstrate genetic divergence, hybridization, and multiple polyploidization in the evolution of Achillea (Asteraceae – Antemideae ). New phytol 166: 273-289.
26- Halliday G., and Beadle M. (1980) Flora Europaea. Volume: 5, Cambridge, London.
27- Inceer H., and Hayirlioglu-Ayaz S. (2007) Chromosome numbers in the tribe Anthemideae (Asteraceae) from north-east Anatolia. Botanical Journal of the Linnean Society, 153: 203–211.
28- Karamenderes C., and Apaydin S. (2003) Antispasmodic effect of Achillea nobilis L. subsp. sipylea (O. Schwarz) Bässler on the rat isolated duodenum. Journal of Ethenopharmacology 84:175-179.
29- Khaniki G. B. (1995) Chromosome numbers and morphometry in Achillea (Anthemideae, Compositae). Nucleus 38: 104-111.
30- Lawrence W. E. (1947) Chromosome numbers in Achillea L. in relation to geographic distribution. Amer. J. Bot. 34: 538–545.
31- Leitch I. J. and Bennett M. D. (1997) Polyploidy in angiosperms. Trends in Plant Science 12: 470-476.
32- Liove F. (1972) IOPB Chromosome number reports. Taxon 21: 333–346.
33- Liove F. (1973) IOPB Chromosome number reports. Taxon 22: 285–291.
34- Maffei M., Germano F., Doglia G. and Chialva F. (1993) Essential oils, chromosome numbers and karyotypes from Italian Achillea L. species. Part II. J. Ess. Oil Res. 5: 61–67.
35- Magulaev A. J. (1982) The number of chromosomes of the species of Asteraceae, Caryophyllaceae and Plantaginaceae of the North Caucasus. Sci. Rep. Higher. School Biological Science, 11: 74-79.
36- Moore D. M. (1982) Flora Europaea check-list and chromosome index. 423 S., Cambridge University press, Cambridge. UK.
37- Morton J. K. (1981) Chromosome numbers in Compositae from Canada and the U.S.A. Bot J Lin Soci 82:357-368.
38- Mozaffarian V. (2005) Notes on the tribe Anthemideae (Compositae), new species, new records and new combination for Iran. Iran. Jour. Bot., 11: 115-127
39- Mozaffarian V. (2007) A Dictionary of Iranian Plant Names, Farhang Moaser Publishers. Tehran, Iran.
40- Oswiecimska M. (1974) Korrelation zwischen Chromosomenzahl und Prochamazulenen in Achillea von Osteuropa. Planta Med., 25: 389-395.
41- Podlech D. (1986) Compositeae VI. Antemideae. In: Flora Iranica, ed. Rechinger K.H., NO.158., pp 49-71. Skademische Druck-u., Verlagsans Talt,Graz, Austria.
42- Post G. E. (1933) Flora of Syria, Palestina and Sinai. American Press, Beirut 2: 44-47.
43- Rahimmalek M., Sayed Tabatabaei B. E., Arzani A. and Etemadi N. (2009) Assessment of genetic diversity among and within Achillea species using amplified fragment length polymorphism (AFLP). Bioch. Syst. Ecol. 37: 354–361.
44- Saukel J., Anchev M., Guo Y. P., Vitkova A., Nedelcheva A., Goranova A., Konakchiev A., Lambrou M., Nejati S., Rauchensteiner F., Ehrendorfer F. (2004) Comments on the biosystematics of Achillea (Asteraceae –Anthemideae) in Bulgaria. Phytol Balcanica, 9: 361-400.
45- Sheidai M., Azanei N. and Attar F. (2009) New chromosome number and unreduced pollen formation in Achillea species (Asteraceae). Acta Biologica Szegediensis. 53:39-43.
46- Soltis P. S. and Soltis D. E. (2000) The role of genetic and genomic attributes in the success of polyploids. Proc. Natl. Acad. Sci. USA, 97: 7051-7057.
47- Soltis D. E., Soltis P. S. and Tate J. A. (2003) Advances in the study of polyploidy since Plant speciation. New Phytologist, 161: 173–191.
48- Stebbins G. L. (1971) Chromosomal evolution in higher plants. Arnold, London.
49- Tutin T. G., Halliday G. and Beadle M. (1980) Flora Europaea. Vol. 5. Cambridge University Press, Cambridge.
50- Vallès J., Garnatje T., Garcia S., Sanz M. and Korobkov A. A. (2005) Chromosome numbers in the tribes Anthemideae and Inuleae (Asteraceae). Kazakhstan. Bot. J. Lin. Soc. 148: 77-85.
51- Watanabe K. (2002) Index to chromosome numbers in Asteraceae. http://www-asteraceae.cla.kobe-u.ac.jp/ index.html.
52- Wendel J. and Doyle J. (2005) Polyploidy and evolution in plants. In: Henry R J (ed.) Plant diversity and evolution: genotypic and phenotypic variation in higher plants. pp. 97-117. Wallingford, CABI Press.
53- Wendel J. F. (2000) Genome evolution in polyploids. Plant Molecular Biology 42: 225–249
54- Wood T. E., Takebayashi N., Barker M. S., Mayrose I., Greenspoon Ph.B. and Rieseberg L. H. (2009) The frequency of polyploid speciation in vascular plants. Proc Natl Acad Sci USA106: 13875-13879.
55- Zargari A. (1996) Medicinal Plants. Tehran University Publication; 4th ed. Vol. 3,. pp. 106-117, Tehran, Iran.
56- Zheng-Yi W., and Raven P. H. (1994) Flora of China. Vol. 17. Science Press, Missouri Botanical Garden, USA.