Amiri, M.J., Eslamian, S.S., 2010. Investigation of Climate Change in Iran. Journal of Environmental
Science and Technology 3, 208–216.
Astrin, J.J., Huber, B.A., Misof, B., Klütsch, C.F.C., 2006. Molecular taxonomy in pholcid spiders
(Pholcidae, Araneae): Evaluation of species identification methods using CO1 and 16S rRNA. Zoologica
Scripta 35, 441–457.
Barrett, R.D.H, Hebert, P.D.N., 2005. Identifying spiders through DNA barcodes. Canadian Journal of
Zoology 83, 481–491.
Darriba, D., Taboada, G. L., Doallo, R., & Posada, D. (2012). jModelTest 2: more models, new heuristics
and parallel computing. Nature methods, 9(8), 772-772.
Dimitrov, D., Astrin, J.J., Huber, B.A., 2013. Pholcid spider molecular systematics revisited, with new
insights into the biogeography and the evolution of the group. Cladistics 29, 132–146.
Esmaeili, H.R., Masoudi, M., ebrahimi, M., Elmi, A., 2016. Review of Aphanius farsicus: a critically
endangered species (Teleostei: Cyprinodontidae) in Iran. Iranian Journal of Ichthyology 3, 1-18.
Frisch, J., 2006. The genus Scopaeus (Coleoptera, Staphylinidae, Paederinae) in Iran, with description of
new species from the Zagros Mountains. Deutsche Entomologische Zeitschrift 53, 5-22.
Gholamifard, A., 2011. Endemism in the reptile fauna of Iran. Iranian Journal Of Animal Biosystematics
7(1).
Hall, T.A., 1999. BioEdit: a user-friendly biological sequence alignment editor and analysis program for
Windows 95/98/NT, in: Nucleic Acids Symposium Series. [London]: Information Retrieval Ltd., pp. 95–
98.
Huber, B.A., 2005. The Pholcid Spiders of Africa (Araneae: Pholcidae): State of Knowledge and
Directions for Future Research, in: African Biodiversity. Springer US, Boston, MA, pp. 181–186.
Huber, B.A., 2011. Phylogeny and classification of Pholcidae (Araneae): an update. The Journal of
Arachnology 39, 211–222.
20 IRANIAN JOURNAL OF ANIMAL BIOSYSTEMATICS Vol.19, No.1
Huber, B.A., 2013. Revision and cladistic analysis of Pholcus and closely related taxa (Araneae,
Pholcidae). Zootaxa 3713, 1–160.
Huber, B.A., Dimitrov, D., 2014. Slow genital and genetic but rapid non-genital and ecological
differentiation in a pair of spider species (Araneae, Pholcidae). Zoologischer Anzeiger-A Journal of
Comparative Zoology 253, 394–403.
Huber, B.A., Eberle, J., Dimitrov, D., 2018. The phylogeny of pholcid spiders: A critical evaluation of
relationships suggested by molecular data (Araneae, Pholcidae). Zookeys 789, 51–101.
Kazemi, S.M. and Hosseinzadeh, M.S., 2020. High diversity and endemism of herpetofauna in the Zagros
Mountains. Ecopersia, 8(4), pp.221-229.
Leigh, J.W., Bryant, D., 2015. POPART: Full-feature software for haplotype network construction.
Methods in Ecology and Evolution 6, 1110–1116.
Mirshamsi, O., 2005. Faunistic study of spiders in Khorasan Province, Iran (Arachnida: Araneae). Iranian
Journal of Animal Biosystematics, 1(1).
Mittermeier, R.A., Robles-Gil, P., Hoffmann, M., Pilgrim, J.D., Brooks, T.M., Mittermeier, C.G.,
Lamoreux, J.L. and Fonseca, G., 2005. Hotspots Revisited: Earth’s Biologically Richest and Most
Endangered Ecoregions (Cemex, Mexico City).
Noroozi, J., Akhani, H., Breckle, S.W., 2008. Biodiversity and phytogeography of the alpine flora of Iran.
Biodiversity and Conservation 17, 493-521.
Noroozi, J., Talebi, A., Doostmohammadi, M., Rumpf, S.B., Linder, H.P., Schneeweiss, G.M., 2018.
Hotspots within a global biodiversity hotspot-areas of endemism are associated with high mountain
ranges. Scientific reports 8, 1-10.
Roewer, C.F. 1955. Die Araneen der osterreichischen Iran-Expedition 1949/50. Sitzungsberichte der
Österreichischen Akademie der Wissenschaften (I) 164, 751-782.
Ronquist, F., Huelsenbeck, J.P., 2003. MrBayes 3: Bayesian phylogenetic inference under mixed models.
Bioinformatics 19, 1572–1574.
Senglet, A., 1974. Pholcus nouveaux d’Iran (Araneae: Pholcidae). Revue suisse de zoologie 81, 803–812.
Senglet, A., 2008. New species of Pholcus and Spermophora (Pholcidae, Araneae) from Iran and
Afghanistan, with notes on mating mechanisms. Revue suisse de Zoologie 115, 355–376.
Silvestro, D., Michalak, I., 2012. RaxmlGUI: A graphical front-end for RAxML. Organisms Diversity &
Evolution 12, 335–337.
Simon, C., Frati, F., Beckenbach, A., Crespi, B., Liu, H., Flook, P., 1994. Evolution, weighting, and
phylogenetic utility of mitochondrial gene sequences and a compilation of conserved polymerase chain
reaction primers. Annals of the entomological Society of America 87, 651–701.
TAXONOMY OF THE GENUS PHOLCUS IN IRAN 21
Wang, Z.L., Yang, X.Q., Wang, T.Z., Yu, X., 2018. Assessing the effectiveness of mitochondrial COI
and 16S rRNA genes for DNA barcoding of farmland spiders in China. Mitochondrial DNA Part A 29,
695–702.
World Spider Catalog, 2021. World Spider Catalog. Version 22.0. Natural History Museum Bern, online
at http://wsc.nmbe.ch, accessed on (Dec 16, 2020)
Zamani, A., Mirshamsi, O., Marusik, Y.M., Moradmand, M. (2021). The Checklist of the Spiders of Iran.
Version 2021, Online at http://www.spiders.ir